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Macro-Moths in Glendaloughin, County Waterford

 

Stiofán MacAmhalghaidh

 

 

Introduction

 

The publication in 2001 of The Lepidoptera of Waterford City & County (LWCC) 1provided the main impetus for the following study. That publication is a catalogue of all lepidoptera recorded in the county of Waterford to the end of 2000, and the first time a single record has been compiled for lepidoptera for this county as a whole. As such, it stands as a landmark publication for lepidopterists in Waterford. Crucially, all records are linked to the 10km squares of the national grid in which they were collected. One fact which becomes very clear when examining records in this way is the contrast between areas with resident recorders and those with none. In particular, those 10km squares with low population, and especially those composed mainly of upland habitats have fared very badly to date. As a resident of the townland of Glendaloughin in square S21, the present author paid particular attention to records from this area. Composed in the main of the slopes of the Comeragh Mountains, S21 is one of those squares with a dearth of records, especially of moths. This is understandable as a large section of S21 is either very lightly populated or without human residents. The low population and inaccessibility of much of S21 aside, O' Meara, in the Introduction in LWCC, notes several factors which tend to hinder the recording of moths in more remote locations:

 

Moths, which are largely nocturnal, must first be caught for examination and identification. The accepted method of doing this is by using modern moth traps that emanate either mercury-vapour or ultra-violet light which attract the moths and catch them alive for examination and later release. If the traps are used in places remote from a supply of electricity, a small portable generator must be used. These are all expensive pieces of equipment, which are not generally available except to the most dedicated fieldworkers.

 

The challenges for the moth enthusiast are not inconsiderable. The problem was thus framed: how to record small, airborne, nocturnal insects in a remote location without excessive expenditure.

 

The solution settled on for the purposes for this experiment was to fit a 25 watt fluorescent tube beside an outdoor white-painted wall and to collect using glass jars which had been stored for recycling. A number of compromises were therefore involved: The lighted wall faces onto a yard enclosed on all sides by farm and domestic buildings, limiting the visibility of the light; recording was limited to a single location; the attractiveness of fluorescent light is not as potent as that of either mercury vapour or ultra-violet lamps; and trapping using individual jars rather than a moth trap necessitates repeated night-time visits by the recorder. These limitations inevitably impact on the number and range of recordings possible, but do at least show what is possible with minimal financial outlay. The issue of record range is mitigated greatly by recording daily rather than on two or three occasions a month. It is hoped that the results detailed below will encourage others across Ireland - and rural Waterford especially - to begin recording.

 

 Waterford ©Michael O Meara MMV

 

Environment

 

Samples were collected from a single location, 200m OD, in the Comeragh Mountains. To the south-east and north-west the land slopes steeply upward, with a small river running north-east to south-west.. The surrounding land-cover includes cattle and sheep pasture and several abandoned fields with a heavy cover of bracken, bramble and, in some places, ragwort. While the immediate vicinity is well-protected and south-facing with a relatively mild micro-climate, within 100m to the north-west the land is very exposed and, where not actively farmed, is a mix of willow, birch, gorse and hazel scrub, and heather-covered upland. Numerous springs allow for several marshy patches within a general context of good, well-drained soils. Also present in the immediate area are considerable patches of several nettle species, several thistle species, an excellent diversity of grasses, and foxgloves, dandelions, wild parsnip, honeysuckle, bilberry, raspberry, blackthorn and other potential larval foodplants. The diversity tree coverage is quite good and includes ash, hawthorn, holly, downy birch, silver birch, crab apple, wild cherry, alder, hazel, goat willow, wych elm, Scots pine and rowan. In addition, several naturalised species are present: English elm, sycamore, beech, white willow, crack willow, and cherry laurel. A plantation of sitka spruce of c.30ha. lies immediately adjacent to the collection site on the eastern side.

 

 

Method of Recording

 

Moths were collected on a nightly basis throughout the period 22 May - 22 September 2005, with the exception of the week 17 - 23 August, on an ongoing basis from dusk through to between 3 and 5am. Specimens were refrigerated to facilitate handling during identification. Identification was by reference to the recently-published Waring and Townsend Field Guide (W&T) 2. In order to limit multiple records of individuals, each was stored until after collecting was completed on the following night and then released approximately 200m from the collection site. Though some micro-moths were recorded where easily identified, the study proper was restricted to macro-moths due to the difficulty of identification of many micro-moth species, and thus no micro-moths have been included in the records given below in Appendix 1. In addition, no records were retained for individuals where identification was in any way uncertain. It is likely that several interesting specimens have not been recorded as a result, though this is considered to have been preferable to including any doubtful records.

 

 

The Records

 

As expected, the quantity of captured individuals on any given night was low, varying from 0 - 28. Almost all were species regularly attracted to light, and most were common species. However, the limited records for S21 to date mean that many otherwise common species are deserving of attention. Rather than discuss each species, therefore, attention in the following four sections focuses on:

· Immigrant species

· Species recorded outside their usual flight season

· Species not previously recorded for County Waterford

· Species not recorded for S21 in LWCC

 

These are followed by a further section of observations arising from the study.

 

 

Immigrant species

 

Only one immigrant species was recorded.

 

Macroglossum stellatarum, Hummingbird Hawkmoth, 22/05

Recorded, on 22 May at mid-afternoon trapped accidentally inside the author's home. This occasion prompted the start of the present study on 22 May rather than the planned date of 1 June. Sightings of this species generally occur later in the summer and, though an arrival in late May is not exceptional, it is not impossible that this individual arrived late in 2004 and overwintered.

 

Macroglossum stellatarum, Hummingbird Hawkmoth

©Michael O Meara MMV

 

Species recorded outside their usual flight season

 

Most individuals in this section occurred only slightly outside the expected season, i.e. by one or two weeks. However, as any such variations are potentially of interest, all such cases are noted.

 

Xanthorhoe designata, Flame Carpet, 07/07, 09/07, 10/07, 13/07

W&T give the flight season as May-June and late July-August, and that for Scotland as June-July. It seems improbable that a single generation as in Scotland applies in Co. Waterford. This is borne out by eight specimens trapped during late August after a month-long gap from mid-July to mid-August. These four specimens, then, represent the first generation and as such are late by 1-2 weeks. This is paralleled by the second generation trapping dates.

 

Eulithis populata, Northern Spinach, 06/07 - 10/09

W&T give flight season as July-August, thus this study seems to show the whole flight season pushed forward about a week, with a dozen specimens appearing during early September. Notable are the ten appearing in the week 04 - 11 September. These constitute the largest concentration in one week for this species over the whole period of the study. Many were worn or weak and may have been attracted to the light as some type of response to weakness. They were easily trapped and perched passively rather than flying.

 

Hydriomena furcata, July Highflyer, 08/07 - 13/09

W&T give the flight season for southern Britain as July-August, and for northern Scotland as late July-early October. It is difficult to state which season the above dates best match, though they seem to parallel the southern British dates, albeit with a late termination in mid-September.

 

Mythimna pallens, Common Wainscot, 24/05 - 29/06

W&T give the first generation flight season in the south of England as June-July. These dates are therefore slightly early but not remarkable. The first specimen was trapped in the first week of the study and the early termination of this generation may have been paralleled by a similarly early start. A second generation was expected, but no evidence for this was found in this study.

 

Amphipyra pyramidea, Copper Underwing, 30/06, 01/07, 04/07, 06/09

W&T give the flight season as starting in late July, thus making these several weeks early. Two specimens were trapped on successive nights, and another three days later. No further specimens were trapped until early September, suggesting two generations, though there seems to be evidence for only one generation elsewhere.

 

Colocasia coryli, Nut-tree Tussock, 27/06, 28/06, 30/06

W&T give the flight season for the first generation in southern Britain as ending in mid-June, thus this about a week late. Six specimens were trapped (three on 28/06), thus this is not a case of a single late specimen.

 

 

Species not previously recorded for County Waterford

 

The reference for this section is LWCC, W&T and personal communication with the author of LWCC, Michael O'Meara.

 

Perizoma blandiata blandiata, Pretty Pinion, 04/07

Trapped at dusk. In Ireland, found mainly in the Burren, County Clare with additional records from widely scattered sites from the south-west to Northern Ireland. No specimen of this species further west than County Cork has previously been recorded in Ireland. As the study site is in west County Waterford, it is possible this individual is a vagrant. It should be noted that this identification has not been verified, though the author has chosen to include this record where several for individuals of other species were not as identification seems clear and the appearance of a vagrant at this location is not outside the range of possibility.

 

Perizoma blandiata blandiata, Pretty Pinion

©Michael O Meara MMV

 

Species not recorded for S21 in LWCC

 

The records for moths in LWCC cover the period up to 2000. Inevitably during the intervening years, species new to the county have been recorded, and species have been recorded from new locations in the county. The effect is that the data in this section is inevitably already out of date. The data will be compared to records for 2000-2005 and when completed, this section will be updated accordingly. As a mark of the very low recording levels for S21, only a handful of species recorded in this study have already been recorded for this square. Species are listed below by English checklist number. A table of all records from this study can be accessed via a link in the Appendix below.

 

Hepialus hecta, Gold Swift

Four specimens from late June and early July. Though apparently very sparsely recorded in Ireland, records are widespread in the county, including the adjacent squares S10, S22 and S30.

 

Hepialus fusconebulosa, Map-winged Swift

Two males. Otherwise sparsely recorded, though the most recent records are from Minella Hotel, Clonmel in 1973, 1974 and Mahon Falls in 1997, both sites in adjacent squares.

 

Falcaria lacertinaria, Scalloped Hook-tip

Four trapped in early August. Previously recorded for Waterford at Clonmel in the early 1900s.

 

Cilix glaucata, Chinese Character

A single specimen was trapped just after dusk in late August. This beautiful and usually quite common moth is represented by four specimens in the county prior to 2000: Clonmel (1947, 1974), Ballymacaw (1978), and Kilmacthomas (1984).

 

Thyatira batis, Peach Blossom

Eleven specimens. Otherwise sparse but widespread records for Waterford include adjacent squares S10 and S22.

 

Habrosyne pyritoides, Buff Arches

Three records, all in late June. Nearest records are for Clonmel in S22. Widely dispersed across the county.

 

Geometra papilionaria, Large Emerald

Ten records. Though best efforts were made to avoid duplicates, the actual number of different individuals trapped was probably 5-8 rather than ten. Otherwise sparsely recorded in Waterford, though recorded by Murray in S22.

 

Hemithea aestivaria, Common Emerald

Twelve specimens recorded from 03 July - 27 July. Records from Clonmel, S22 exist from the 1930s, 1960s and 1970s, but all other records for the county are from the south east and south west.

 

Scopula immutata, Lesser Cream Wave

A single specimen in early July. Sparse but widespread records for the county. The most recent in 1986 near Cappoquin, but also from Clonmel in the 1920s and '30s.

 

Scopula floslactata, Cream Wave

Three specimens trapped within a single week in early July. Previous records are sparse, from the area of Waterford City in 1997, and from Clonmel in the Murray collection.

 

Xanthorhoe designata, Flame Carpet

Twelve specimens recorded during July, August and early September. First recorded for the county in 1974 in Clonmel. Current records are from three adjacent squares: S11, S22, S30 and from Waterford Airport. There is clearly a population resident in the Comeraghs south of Clonmel.

 

Xanthorhoe ferrugata, Dark-barred Twin-spot Carpet

Seven specimens trapped over three weeks from late July to mid-August. Previously recorded only at Clonmel in the 1920s and '30s, and in 1974, and in 1998 in Waterford City. The numbers suggest that a population is resident in the Comeraghs just south of Clonmel accounting for the records in this study and earlier records from Clonmel.

 

Xanthorhoe montanata montanata, Silver-ground Carpet

Twenty-three trapped throughout June. Other records are focused on the eastern end of the county, and on Clonmel.

 

Scotopteryx chenpodiata, Shaded Broad-bar

A single specimen trapped in early August. Previously recorded in several sites in east Waterford and at Clonmel.

 

Epirrhoe alternata, Common Carpet

One trapped in mid-August, with two more in early September. Previously recorded from almost every other area of the county, including four adjacent squares.

 

Camptogramma bilineata bilineata, Yellow Shell

A single specimen recorded in early July, disturbed from shrubs while walking. Records exist from across the county though only those from Clonmel are adjacent.

 

Entephria caesiata, Grey Mountain Carpet

A single specimen from the start of August. Previously only known for the county from Clonmel, in the Murray collection.

 

Mesoleuca albicillata, Beautiful Carpet

Two specimens were trapped in mid-August. The nearest record from Co Waterford comes from S22.

 

Cosmorhoe ocellata, Purple Bar

One trapped in mid-September. Otherwise only recorded at Clonmel and Killahaly Wood.

 

Eulithis populata, Northern Spinach

Twenty-eight specimens trapped over nine weeks. Very sparse records previous to this significantly include Clonmel, less than ten miles away.

 

Ecliptopera silaceata, Small Phoenix

Five records during August. Not recorded for Waterford County in LWCC, though it may have been identified since. This is currently being checked.

 

Chloroclysta siterata, Red-green Carpet

A single specimen. Previously recorded only at Clonmel by Murray.

 

Chloroclysta citrata, Dark Marbled Carpet

Twenty-one specimens recorded between 15/07 - 28/08. Common where active recorders present, suggesting it is very under-recorded. The present study supports this. Recorded from S11 and S22.

 

Chloroclysta truncata, Common Marbled Carpet

Thirty-four specimens trapped between 14/08 - 21/09. Common across the county and recorded from three adjacent squares: S11, S22 and S31

 

Colostygia pectinataria, Green Carpet

Twenty-three trapped during May and June. Also recorded from adjacent squares S22 and S30.

 

Hydriomena furcata, July Highflyer

Numerous records through July, August and September. Well-recorded elsewhere in the county.

 

Euphyia unangulata, Sharp-angled Carpet

Twenty trapped during July and August, though the inclination of individuals to return to light the night after release may mean the actual number of individuals is about half that stated. Previous records for the county are widespread and include S22 and S30, both adjacent.

 

Perizoma taeniata, Barred Carpet

Two specimens were trapped in early August. Though there is no record for this species in LWCC, specimens may have been found since 2000. This is currently being confirmed.

 

Perizoma blandiata blandiata, Pretty Pinion

A single record. Details given for this species in the previous section as it is otherwise unrecorded for the county.

 

Perizoma didymata didymata, Twin-spot Carpet

A single specimen at the start of September. Previous records are from Clonmel, and one from Ballymacaw in 1977.

 

Eupithecia tenuiata, Slender Pug

A single record from the start of August. One other record, from 1901, completes the county inventory.

 

Eupithecia absinthiata, Wormwood Pug

Two specimens trapped at the start of August. Murray recorded the species at Clonmel, and one other record exists from Waterford City, 1979.

 

Eupithecia vulgata, Common Pug

Two specimens trapped in early and late June. Also recorded for adjacent squares S22 and S31.

 

Eupithecia tripunctaria, White-spotted Pug

A single specimen trapped in mid-August. Other county records are from Clonmel, S22 in the early 1900s, and Kilbarry Bog, S61 in 1997.

 

Gymnoscelis rufifasciata, Double-striped Pug

Three specimens recorded in a single week. Other records exist from across the county, including adjacent S22.

 

Opisthograptis luteolata, Brimstone Moth

This species is widely distributed in the county, with the nearest records from S22. Six specimens from the end of July through to late September.

 

Epione repandria, Bordered Beauty

The nearest records are from S22. The number of specimens trapped - ten, all in the month of August - suggest that this species is not uncommon in the area.

 

Aperia syringaria, Lilac Beauty

Four trapped in a fortnight around the start of July. Previously, sparse records from the early 1900s constitute the whole record for the county, though significantly including records from S22.

 

Ennomos alniara, Canary-shouldered Thorn

A single specimen was trapped in late August. Previously recorded at three sites in the county, the nearest at Clonmel, S22.

 

Selenia dentaria, Early Thorn

Fifteen specimens trapped in late July and through August. Sparsely recorded elsewhere, but including S22.

 

Biston betularia, Peppered Moth

Eight records from May-July. Sparse but widespread records for the county include Clonmel to the immediate north in S22.

 

Biston betularia, Peppered Moth

©Michael O Meara MMV

Cabera exanthemata, Common Wave

Seven records throughout July. Recorded from three other locations in Waterford including adjacent S22.

 

Campaea margaritata, Light Emerald

Ten trapped between late June and mid-July. Previously only recorded at Clonmel in the 1930s and 1960s, and more recently in the extreme south-west of the county.

 

Laothoe populi, Poplar Hawkmoth

Two records in one week may represent a single individual, though the area is well populated with larval foodplants. Both specimens were trapped shortly before dawn. Otherwise widespread, sparse records may represent poor recording.

 

Phalera bycephala, Buff-tip

A single record from early July. Only recorded four times for the county according to LWCC, most recently at Waterford City. The nearest record is also the earliest, at Clonmel in 1974. The previous records are widely dispersed across the county.

 

Phalera bycephala, Buff-tip larva

©Michael O Meara MMV

Ptilodon capucina, Coxcomb Prominent

A single specimen from early July. Previously only recorded from Youghal in 1981 and Killahaly Wood in 1986.

 

Thumatha senex, Round-winged Muslin

Two trapped in early July. It is worth quoting O'Meara in full on this rare species: "There appears to be more records of this rare moth for Waterford than for the whole of the rest of Ireland. Dr. H. Murray found it at Cappagh at the end of June 1931 and Baynes (1964) quotes this as being one of only three records for Ireland. In addition, de Worms (1983) gives a more recent record for south of Waterford City while in July 1997, one which flew to a lighted window in the City was later determined by K. G. M. Bond (pers. comm.) as this species. De Worms loc. cit. gives only two other known locations in Ireland." (LWCC, 62)

 

Nudaria mundana, Muslin Footman

A single specimen recorded in July. Restricted almost wholly to west Cork and Kerry, two records for Waterford exist, at Bunmahon in 1997 and Mt Congreve in 2000. It may be that the south-west population is spreading eastwards, though the species may simply be poorly recorded in the area to date.

 

Lithosia quadra, Four-spotted Footman

A single specimen trapped in early August. Previous records for the county end with a record by Murray in the early 1900s at Clonmel.

 

Spilosoma lubricipeda, White Ermine

71 specimens trapped between late May and late July. The period 29/05 - 18/06 was highly productive, giving 51 of the total. Common and widespread in the county. Nearest records are for Clonmel, S22.

 

Spilosoma lutea, Buff Ermine

Nine specimens trapped during late June and early July. Common and widespread in the county. Nearest records are for Clonmel, S22.

 

Euxoa nigricans, Garden Dart

Eight records in July and August. Previously recorded only in the vicinity of Waterford City.

 

Ochropleura plecta, Flame Shoulder

Three specimens in early July. Widespread and common in the county. Nearby records exist from S22 and S30.

 

Noctua pronuba, Large Yellow Underwing

A single specimen from early July. Of this species, LWCC says: "The Large Yellow Underwing is probably the most common moth in the county and up to two hundred have been recorded in moth traps on many occasions." (LWCC, 68) Despite this, it is previously unrecorded for S21, highlighting the sparsity of records for this 10km square.

 

Noctua comes, Lesser Yellow Underwing

A single specimen trapped in mid-August. Widespread in the county including two adjacent squares.

 

Noctua janthe, Lesser Broad-bordered Yellow Underwing

A single specimen. Otherwise widespread in the county.

 

Lycophotia porphyrea, True Lover's-knot

A single specimen recorded in mid-June. Otherwise widespread and quite common across the county. Previous records include S22 and S30, both adjacent squares.

 

Diarsia brunnea, Purple Clay

A single specimen trapped in early July. The only other Waterford records are three specimens from Clonmel, Killahaly Wood and the vicinity of Tallow.

 

Xestia baja, Dotted Clay

Recorded in late July-early August. Only one other record, from the Tallowbridge area in W99 (which is only partly in Waterford), may also be from the county.

 

Ceramica pisi, Broom Moth

Not recorded for the county before 1984, several have since been seen, with the nearest records from just south of Mahon Falls in the adjacent square, S30. Eight records from the end of June through to mid-July.

 

Mythimna pallens, Common Wainscot

Recorded continuously from the start of the study through to the end of June. Widespread across the county including three adjacent squares: S10, S22, S31.

 

Blepharita adusta, Dark Brocade

One specimen from early July. For this species, LWCC states: "Several to light at Ballymacaw in June 1975 is the only record for Waterford." (LWCC, 76)

 

Amphipyra pyramidea, Copper Underwing

Five specimens from late June/early July and early September. Previously only recorded from Clonmel, S22.

 

Euplexia lucipara, Small Angle-shades

A single record for early July. Previous records are from widely spaced sites across the county, including Clonmel.

 

Phlogophora meticulosa, Angle Shades

Four specimens were trapped between late August and mid-September. The nearest record from Co Waterford comes from S22.

 

Mesapamea secalis, Common Rustic

Eight specimens trapped between early and late August. Recorded from widespread locations across the county including adjacent squares S10 and S31.

 

Chortodes pygmina, Small Wainscot

Fourteen specimens trapped between late August and early September. Common in boggy areas, though recorded in LWCC only around Waterford City, in coastal squares and at Clonmel.

 

Gortyna flavago, Frosted Orange

A single specimen trapped in mid-September. Few records for the county, and the most recent dates to 1964.

 

Coenobia rufa, Small Rufous

Three specimens trapped, two in early August, one in late August. A rare species listed for just nine locations in Ireland in 1983, one near Waterford City.

 

Colocasia coryli, Nut-tree Tussock

Twenty-five specimens, most first generation. Previously recorded only at Clonmel in the early 1900s, and seven from Kilbunny Wood near Portlaw in 1997.

 

Autographa pulchina, Beautiful Golden Y

O'Meara notes "This is not a common moth in Waterford and when found is usually in small numbers." (LWCC, 84) and this is borne out by the two specimens in this study, found in early August. The nearest previous record in LWCC is for S22.

 

Abrostola tripartita, Spectacle

Three recorded in a single week. Records are widespread in the county, though locations are few.

 

Scoliopterix libatrix, The Herald

A single specimen trapped in mid-September. Recorded intermittently for the county including at Clonmel.

 

Hypena proboscidalis, Snout

Numerous records throughout July and August. Common and widespread including three adjacent squares: S10, S22 and S30

 

Schrankia costaestrigalis, Pinion-streaked Snout

Numerous records throughout July and August stand in stark contrast to just two previous recorded individuals for the county, one for Murray at Clonmel in 1935, one for Kilmacthomas. Both previous records, however, occur in squares adjacent to S21 and at opposite sides of it, suggesting that a population may exist centred in S21, with previous records being made at the extremes of the population's range.

 

Zanclognatha tarsipennalis, Fan-foot

Thirteen specimens from late June to mid-July. Previous records for the county are from Clonmel in adjacent S22.

 

Herminia grisealis, Small Fan-foot

Six trapped during late June and early July. Previously recorded from Clonmel and Waterford City.

 

 

Observations

 

The minimal cost with which this study was carried out demonstrates that such recording work is feasible on an ongoing basis. The low numbers recorded each night ensure that the burden of identification is light, allowing the amateur time to take care over identification. By sampling small numbers on an ongoing nightly basis, a wholly different picture of local populations is produced from that obtained by intensive study over a short period. While the latter method produces larger numbers of specimens for a given date, unfortunate timing can easily result in important species not being observed at all. The main problem in using the method employed in this study lies in the need to attend the study site on an ongoing basis during the night. There is no easy solution to this, though replacement of the current light source with a home-made moth trap, which can be built at minimal cost and can be designed to be portable, will remove the need to tramp about in darkness night after night.

 

The section above titled Species not recorded for S21 in LWCC demonstrates the dearth of records in the county for the remoter 10km squares, and also demonstrates clearly the impact even casual recording can have on the county record. Only a few species recorded by this study had already been identified in S21. Seventy-seven species recorded are new to S21. One species is new to the county, and two others are possibly new to the county. While it is unlikely that this level of success will ever be matched in S21 again, there is every reason to expect similar levels of success for new recorders located in similar remote sites in the county.

 

The possibilities of using a portable lamp or trap in 2006 are now being considered with a view to expanding the geographical range of recording in the area.

 

 

Appendix

 

A table listing all species by week for the study period can be viewed or downloaded by clicking here.

 

 

References

 

(1) Michael O'Meara, The Lepidoptera of Waterford City & County, Waterford Wildlife, 2001

 

(2) Paul Waring and Martin Townsend, Field Guide to the Moths of Great Britain and Ireland, British Wildlife Publishing, 2003

 

 © Stiof MacAmhalghaidh MMV

 

About the author

 

Stiofán MacAmhalghaidh is currently lecturing part-time in archaeology and history at Waterford Institute of Technology. He founded the IRQUAS Irish Studies Online Project in 2000, and is Editor of both Insight and Celtic Well online journals. Stiof has been photographing and recording fungi, trees, lichens and moths since 1993, and is currently working on a digital encyclopaedia of Irish native tree species.

 

 

 

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